American chestnuts, towering 30 meters or more, once dominated forests throughout the Appalachian Mountains. But in the early 1900s, a fungal infection appeared on trees at the Bronx Zoo in New York City, and then spread rapidly. The so-called chestnut blight—an accidental import from Asia—releases a toxin that girdles trees and kills everything above the infection site, though still-living roots sometimes send up new shoots. By midcentury, large American chestnuts had all but disappeared.
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